Habitat: medium-altitude forest valleys.
BotanyFern; creeping; summer-green.
Root: rhizome; creeping; 2 to 4 mm in diameter; with sparse yellowish brown, broadly lanceolate, thinly membranous scales. Leaves: fertile frond 40 to 90 cm; stipe chestnut-black or purple-brown, 15 to 40 cm, ± 2 mm in diameter, grooved adaxially, with sparse scales at base, upward glabrate, shiny; lamina 2 or 3 pinnate, broadly ovate to deltoid-ovate; pinnae 10 to 15 pairs, opposite or subopposite, 3 to 4 cm apart, spreading or ascending, subsessile or very shortly stalked.
Sori: orbicular; abaxial on basal acroscopic veins; indusia pale green, broadly ovate, small, membranous, glandular, sparsely ciliate at margin, proximal to sorophore, hidden by sporangia at maturity, persistent.
Spores: yellow; bean-shaped; spheroidal, perispore with dense inflated, bulbous processes.