Synonyms; Antigramma cardiophylla; Asplenium cardiophyllum; Boniniella cardiophylla; Phyllitis cardiophylla.
Clades: Hymenasplenioideae;
Aspleniaceae;
Aspleniidae;
Pteridophyta;
Plants.
Genus: ± 35 species.
Region: Vietnam, China, Japan.
Habitat: subtropical or tropical moist lowland forests; on rocks or sandy soils, in forests, near streams.
Ecology: threatened by habitat loss.
BotanyFern; sexual; diploid; up to 30 cm tall.
Root: rhizomes long, creeping, with blackish brown scales, with yellowish brown hairs; ovate-triangular, small, margins sparsely toothed, and caducous on older rhizome parts.
Leaves: well separated; simple; thinly papery; dark to brownish green when dry; subglabrous; stipe shiny, dark brown to black, 10-20 cm, terete but with adaxial groove, base with scales and hairs; lamina simple, ovate, 9 to 14 by 5 to 9 cm, base cordate, margin entire or shallowly sinuate, apex acute to acuminate; rachis obvious abaxially, shiny and black to middle of leaf, lateral veins anadromous, slender, and hardly visible on adaxial side, occasionally connected and forming elongate areoles near margin, vein ends free.
Sori: linear; usually solitary; on acroscopic veins; rarely opposite; indusia persistent; brownish; thinly membranous; entire.
Spores: elliptic; 21 to 24 µm long; perispore with narrow crests.