Name: from the botanist Antoine Fée.
Synonyms: Grammitis vulcanica; Pleopeltis feei.
Indonesia: pakis tangkur.
Region: Indonesia, Sumatra, Java, Lesser Sunda Islands; China, Philippines, Indonesia, Malaysia, Pacific Islands.
Habitat: terrestrial; tropical, subtropical; monsoonal; soil moist, fertile, loamy; forest, open heath, between rocks, on cliffs and roadsides; elevation 900 to 3150 m; seldom on walls or epiphytically on mossy trunks; especially on volcanoes, near craters, resistant to volcanic fumes; prefers a coarse mixture, enough shade, humidity, air movement.
Content: selligueain A, selligueain B, (-)-4-β-carboxymethyl-epiafzelechin-(3'-deoxydryopteric acid), (+)-afzelechin-O-β-4'-D-gluco-pyranoside and kaempferol-3-O-β-D-glucopyranoside-7-O-α-L-rhamnopyranoside; selligueain, a proanthocyanidin trimer and a sweetener; kaempferol-3-O-β-D-glucopyranoside-7-O-α-L-rhamnopyranoside, a known bitter-tasting flavonoid glycoside: (-)-4β-carboxymethyl epiafzelechin (3'-deoxydryopteric acid); epiafzelechin-(4β→8, 2β→O→7)-epiafzelechin-(4β→8)-3'-deoxydryopteric acid methyl ester (selligueain B); (+)-afzelechin-O-β-4'-D-glucopyranoside.
Use: ornamental, parks, gardens, in pots and baskets.
BotanyFern; evergreen; epiphyte, lithophyte; wide creeping.
Root: rhizome, short, creeping, 3 to 8 mm diameter, densely covered with peltate, rarely pseudopeltate scales; scales lanceolate to linear, ± 6 mm long, ± 2 mm wide, obtuse or more often acute, golden-brown, evenly coloured, entire to rarely remotely and weakly dentate, appressed or more often spreading.
Stem: densely scaly when young; scales are reddish-brown.
Leaves: simple; dimorphous; stipe of sterile fronds are 18 to 20 cm long, lamina measuring about 15 to 20 by 7 to 9 cm; leathery; glabrous; stipe of fertile fronds measuring about 20 to 35 cm long, lamina ovate and measuring about 13 to 20 by 3 cm.
Leaves: distance between leaves 1-4.5 cm; vascular strands 10 to 13, with bundle sheath fully sclerified, no sclerenchyma strands.
Fertile leaves: petiole firm, 4 to 55 cm long, glabrous, erect, lustrous pale or brown; lamina lanceolate, 7 to 25 cm long, 2 to 7 cm wide; base obtuse, margin cartilaginous, thickened; without or with few notches; apex various, obtuse to acuminate; lustrous bright green; coriaceous; glabrous.
Sterile leaves: usually present, petiole 2 to 45 cm long; lamina ovate-lanceolate, 5 to 31 cm long, 2 to 10 cm wide; main veins on adaxial surface raised or not, distinct; veinlets indistinct, free, anastomosing, excurrent and recurrent; hydathodes frequent, calcareous scales caducous.
Sori: long; linear; between the main veins; reddish-brown;mostly confluent across connecting veins into interrupted transverse linear coenosori, 3 to 5 mm wide; in one row between adjacent costules from the costa to the leaf margin, not quite reaching either the midrib or the margin, superficial.
Spores: densely granulose surface.
Chromosome number: 2n= 74.