English: Fishbone fern; Tuberous sword fern; Tuber ladder fern; Erect sword fern; Narrow sword fern; Ladder fern; Herringbone fern.
Hawaiian: Kupukupu; Okupukupu, Ni'ani'au.
Region: global tropics, northeastern Australia, Asia, Himalayas; naturalised central east coast of New South Wales; introduced into Bermuda, French Polynesia, New Zealand, United States, tropical North, Central and South America, Mexico, Caribbean Florida, Africa, Southeast Asia, Pacific islands, Azores.
Habitat: in moist, shady groves, rock faces, roadsides, on old walls or ruins; in swamps and floodplains in coniferous forests.
BotanyFern; evergreen; 40 to 80 to 100 cm tall; terrestrially, rarely an epiphyte.
Root: underground rhizome in the form of several small tubers.
Leaves: pinnate; erect and pinnate linear to lanceolate, glandular and simple; rachis bears bicolored chaff scales; petiole covered with bicolored pale and dark brown scale; leaflets entire, sessile and elongate-lanceolate; 5 centicmmeters long and up to 1 cm wide, standing at a distance of less than 1 centimeter.
Sori: rounded.
Spores: warty, wrinkled.