Clades:
Dicranidae;
Bryanae;
Bryophyta;
Plants.
BotanyCentral strand absent. Leaves cells (rounded-)quadrate. Costa present. Dioicous. Perichaetia developing after fertilization. Antheridia without specialized capsule. Seta short, massive. Capsule valves irregular (often 4-5 main and 1-2 shorter ones), also separate at apex Spore sac dome-shaped. Germination endo- sporic. Calyptra covering entire capsule.